This is a ternary operator, used as a shorthand conditional statement:
it's the same as saying:
if ($.browser.mozilla) {
mouseWheelEventName = 'DOMMouseScroll';
} else {
mouseWheelEventName = 'mousewheel';
}
The first piece before the = is declaring a variable (mouseWheelEventName) dependent on the following condition.
The next piece between = the ? is the condition (is $.browser.mozilla true?).
Immediately after the ? is the then portion (if the condition is true, set the variable mouseWheelEventName to the string DOMMouseScroll).
After the : is the else (if the condition is NOT true, set the variable mouseWheelEventName to the string mousewheel).
Further reading:
https://developer.mozilla.org/en-US/docs/Web/JavaScript/Reference/Operators/Conditional_Operator
As for why there is a comma at the end of it, we'd need to see a more complete code sample including what follows that to say for certain. If I had to guess, I would say the author of the code was chaining variable statements. This answer may shed some light for you: Javascript best practices, why to use comma to chain function/variable declarations? (see chosen answer)